Page image

polyclonum Haussk., Temple River, W.5. porphyrium Simpson, Olivine Range, W.8. pubens A. Rich., near Opotiki, W.9. var., Mt. Hikurangi, East Cape, R.20. purpuratum Hook. f., head of Matukituki River (West Branch), W.9. pycnostachyum Haussk., Mt. Torlesse, W.14 Mt. Terako, R.14. rostratum Cheesem., Ohau River, N. Otago, R.15. var. pubens Petrie, Molesworth Station, W.6. rotundifolium Forst., Gouland Downs, W.9. tasmanicum Haussk., Mt. Torlesse, W.9. tenuipes Hook. f., Old Man Range, W.7. vernicosum Cheesem., Mt. Peel, N. W. Nelson, R.15. sp. with glandular hairs on leaves, shingle slides, Mt. Terako, W.12. Flower Structure In the New Zealand species the flower structure follows much the same pattern of variation as that occurring among members of the genus in other parts of the world. Size of Flowers The flowers range in size from a diameter of about 4 mm in E. nerterioides to 25 mm in E. chloraefolium var. kaikourense and E. pallidiflorum. Those of 21 of the species and varieties listed measure less than 10 mm diameter; 15 of them are between 10 and 15 mm diameter; 4 are over 15 mm diameter. Flower Colour Thirty have white flowers, 7 are rose coloured and 3 purple. Calyx Limb The 4-partite calyx limb reaches almost as high as the corolla in one species, E. cinereum and, in the other extreme, in one species, E. chloraefolium (Arthur's Pass), it reaches only half way up the corolla. Relative Position of Anthers and Stigma. In all except one species the introrse anthers of at least the four longer of the eight stamens at their dehiscence are level with some part of the stigma, usually the lower portion. After the commencement of their dehiscence the filaments lengthen a little, and in some species, notably E. rotundifolium, some of the anthers stand well above the stigma. The exception is E. chloraefolium var. kaikourense in which all the anthers are always far below the stigma level. In E. chionanthum the four longer anthers at the commencement of their dehiscence are slightly below the receptive part of the stigma but a day later they may overtop it. I have been unable to detect any special advantage deriving from the four short stamens except the negative one that overcrowding of the anthers of the four longer stamens is avoided. In certain circumstances this permits the latter and, in some species, also those of the short stamens when they have dehisced to affix themselves to the stigma in a neat, closed ring. Pollen The mature pollen grains remain attached in tetrads. By tapping them on a glass slide with a penknife the tetrads can be broken into their four pollen grains. Normally they are transferred unbroken to the stigma. Each grain usually bears a delicate, viscous and often curled filament, in length usually two to four times the diameter of the grain. Commonly four of these filaments may be seen to project from a tetrad. They function as grappling hooks and very frequently hold the tetrads together in masses or strung like beads.