Carpellody in the Wheat Flower and Its Inheritance. By J. W. Calder, M.Sc., B.Ag., Canterbury Agricultural College, Lincoln. [Issued separately, 23rd August, 1930.] The term carpellody signifies sex-reversal in the direction of male to female organs in plants. An interesting abnormality of the wheat flower was noticed in a plant among the wheat plots at Lincoln College in 1924. The plant first attracted notice because the glumes remained expanded for from seven to ten days giving the head a conspicuous transparent effect. The abnormality has been investigated and certain experiments have been conducted to determine its behaviour on breeding. The present paper is a progress report of the results. Carpellody has been reported in various plants and consists of partial or complete transformation of stamens to carpel. De Candolle reported carpellody in the wallflower (Cheiranthus cheiri and he gave the plant the varietal name C. cheiri gynanthus. This case has been investigated by Nelson in the Publications of the Royal Society of Tasmania in 1928. The stamens adhere together and form a closed ring round the normal carpel. Ovules are produced within this ring and when pollinated produce viable seeds. Nelson crossed a carpellodic form with a normal. The F1 from the cross were normal and segregation occurred in the F2 in the proportion of 3 normal to 1 carpellodic showing that it was a simple mendelian recessive character. Weatherwax, in the Proceedings of the Indiana Academy in 1925 reported carpellody in Maize, where the rudimentary anthers of the pistillate flowers were transformed into carpelloid structures. In this case, however, he states that there is no true ovarian cavity and no ovules are formed although style and stigma are similar to the normal. Shaffner, in the American Naturalist in 1925, when discussing sex differentiation and determination in higher plants, states that sex-reversal is primarily dependent on physiological states, and these are subject to change and reversal through ecological factors. The wheat flower is considered extremely stable few abnormalities having been reported. Anthony, in the Journal of Heredity, 1918, reports an abnormality in which the anthers bore a few stigmatic hairs and the filament united with the anther as one piece. Leighty and Sando, in the Journal of Heredity in 1923, reported pistillody in the wheat flower and from their description the abnormality is the same as the one here described. They did not give any anatomical description and they state that the character is not