Page image

a depth of 23mm.; so that the anterior width is less than the depth. The anterior surface is not so oblique as is usual in the other small species of moa, but makes almost a rightangle with the outer surface. It is also directed inwards well within the inner edge of the middle trochlea (fig. 4). The most remarkable character, however, is its flatness behind (fig. 5), the posterior process, which is so large in all other moas except Palapteryx dromioides, being here quite small. The trochlear groove is slight. The middle trochlea is considerably broader than the inner one, its width being probably about 28mm., but it tapers off rapidly so that its distal width is only about 19mm. Its depth is 26mm. It does not project so far forwards as in most other moas. The intertrochlear gorge is broad and does not narrow so much at its entrance (figs. 3–4) as in all other moas except Cela curta. Its width at the entrance is nearly half that of the inner trochlea. Supposed Femur. Plate IV., fig. 6. This is nearly cylindrical, the greatest and least diameters being 28mm. and 26mm. It is remarkable for the thickness of the bony wall, the diameter of the medullary cavity being only about 8mm. or 9mm. This great thickness of the bony wall makes it doubtful if the bone belongs to a bird at all. The only bones of the moa which have so relatively small a central cavity are the toe-bones, and these are always flat on the undersurface. It is also too large for a toe-bone of A. antiqua, while its diameter is about what we should expect in the femur of that species. As the bone has disappeared it is impossible now to make a microscopical investigation to test its avian or reptilian character. Affinities of the Species. In the high intercondylar ridge our species resembles Anomalopteryx didiformis, Mesopteryx didina, and Pachyornis (?) geranoides, and differs from Cela curta. In the sinuated anterior margin of the proximal articular surface it most resembles A. didiformis and P. geranoides, but in these species there is usually only a single concavity on the inner side of the intercondylar ridge. In the absence of a longitudinal ridge on the inner margin it resembles P. geranoides but this ridge is often absent in A. didiformis and C. curta. In the high ectocondylar margin it most resembles M. didina and P. geranoides. In the small inner hypotarsal ridge it approaches Emeus crassus, but A. didiformis sometimes has the inner ridge less than the outer.